Scientific publications (see bottom of the page for manuscripts under review):
27. Roth, T. S., Massen, J. J. M., & Samara, I. (in press). Inconsistent Use of Evolutionary Frameworks in Accounts of Romantic Relationships (invited commentary). Behavioral and Brain Sciences.
26. Roth, T. S. (2026). Incorporating free partner choice into conservation breeding: the case for long-distance preference testing. Behaviour, https://doi.org/10.1163/1568539X-bja10346
25. van Leeuwen, E. J. C., & Roth, T. S. (2025). On the forces that bind us (invited commentary). Behavioral and Brain Sciences, 48, e186. https://doi.org/10.1017/S0140525X25101477
24. de Laat, E., Waasdorp, S., Roth, T. S., Massen, J. M. M., & Sterck, E. H. M. Despotic long-tailed macaques benefit others in a group service paradigm. Behaviour, 162(15), 1079–1107. https://doi.org/10.1163/1568539X-bja10338
23. Roth, T. S., van Berlo, E., Perea-García, J. O., & Kret, M. E. (2025). Bornean orangutans (Pongo pygmaeus) show an attentional bias toward a male secondary sexual trait. Annals of the New York Academy of Sciences, 1552, 225-238. http://doi.org/10.1111/nyas.70032
22. Sterck, E. H. M., Kamp, S. M., Rouart, I., van den Berg, L. M., Zijlmans, D. G. M., Roth, T. S., & Bushman, B. J. (2025). Reactions to Social Videos in Long-tailed Macaques. Animal Cognition, 28, 50. https://doi.org/10.1007/s10071-025-01970-1
21. van Oosten, J., Vanhooland, L. C., Roth, T. S., Gallup, A. C., Massen, J. M. M. (2025). No experimental evidence for yawn contagion in galahs (Eolophus roseicapilla). Animal Behavior & Cognition, 12(2), 148-171. https://doi.org/10.26451/abc.12.02.01.2025
20. Benítez, M. E., & Roth, T. S. (2024). Frans de Waal’s lasting legacy: Inspiring a new generation of animal behaviour scholars. Behaviour, 161(7), 511-522. https://doi.org/10.1163/1568539X-bja10277
19. Roth, T. S., von Wulffen, C., Fox, M., Askew, J., Spierings, M., & Perea-Garcia, J. O. (2024). Idiosyncratic long call-like vocalizations in two zoo-housed Bornean orang-utan females. Behaviour, 161(7), 567-585. https://doi.org/10.1163/1568539X-bja10273
18. Roth, T.S., Samara, I., Perea-Garcia, J.O., & Kret, M.E. (2024). No immediate attentional bias towards or choice bias for male secondary sexual characteristics in Bornean orang-utans (Pongo pygmaeus). Scientific Reports, 14, 12095. https://doi.org/10.1038/s41598-024-62187-9
17. Roth, T.S. (2024). Tinder for orang-utans: comparing sexually selective cognition among Bornean orang-utans (Pongo pygmaeus) and humans (Homo sapiens). PhD Thesis, Leiden University. https://hdl.handle.net/1887/3721951
16. Folz, J., Roth, T.S., Nikolic, M., & Kret, M.E. (2024). Who gets caught by the emotion? Attentional biases toward emotional facial expressions and their link to social anxiety and autistic traits. Current Psychology, 43, 3486–3502. https://doi.org/10.1007/s12144-023-04484-6
15. Roth, T.S., Samara, I., Perea-Garcia, J.O., & Kret, M.E. (2023). Individual attractiveness preferences differentially modulate immediate and voluntary attention. Scientific Reports, 13, 2147. https://doi.org/10.1038/s41598-023-29240-5
14. Laméris, D.W., van Berlo, E., Roth, T.S., & Kret, M.E. (2022). No evidence for biased attention towards emotional scenes in Bornean orangutans (Pongo pygmaeus). Affective Science, 3, 772-782. https://doi.org/10.1007/s42761-022-00158-x
13. Samara, I., Roth, T.S., Nikolic, M., Prochazkova, E., & Kret, M.E. (2022). Can third-party observers detect attraction in others based on subtle nonverbal cues? Current Psychology. https://doi.org/10.1007/s12144-022-02927-0
12. Sterck, E.H.M., & Roth, T.S. (2022). Socioecology of Asian colobines. In: The Colobines: Natural History, Behaviour, and Ecological Diversity (Matsuda I, Grueter C & Teichroeb JA, eds.). Cambridge University Press, UK (pp 250-270). https://doi.org/10.1017/9781108347150.017
11. Roth, T.S., Du, X., Samara, I., & Kret, M.E. (2022). Attractiveness modulates attention, but does not enhance gaze cueing. Evolutionary Behavioral Sciences, 16(4), 343-361. https://doi.apa.org/doi/10.1037/ebs0000265
10. Samara, I., Roth, T.S., & Kret, M.E. (2021). The Role of Emotion Projection, Sexual Desire, and Self-Rated Attractiveness in the Sexual Overperception Bias. Archives of Sexual Behavior, 50, 2507–2516. https://doi.org/10.1007/s10508-021-02017-5
9. Nikolic, M., & Roth, T.S. (2021). A comparative glance on self-conscious emotions: A commentary on Kret et al. (2020). Neuroscience & Biobehavioral Reviews, 129, 154-156. https://doi.org/10.1016/j.neubiorev.2021.07.031
8. Roth, T.S., Samara, I., & Kret, M.E. (2021). Multimodal mate choice: Exploring the effects of sight, sound, and scent on partner choice in a speed-date paradigm. Evolution and Human Behavior, 42(5), 461-468. https://doi.org/10.1016/j.evolhumbehav.2021.04.004
7. Roth, T.S., Samara, I., Tan, J., Prochazkova, E., & Kret, M.E. 2021. A comparative framework of inter-individual coordination and pair-bonding. Current Opinion in Behavioral Sciences 39. https://doi.org/10.1016/j.cobeha.2021.03.005
6. Roth, T.S., Samara, I., & Kret, M.E. (2021). Ultimate and proximate factors underlying sexual overperception bias: A reply to Lee et al. (2020). Evolution and Human Behavior, 40(1), 73-75. https://doi.org/10.1016/j.evolhumbehav.2020.06.002
5. Roth, T.S., & Sterck, E.H.M. (2020). Social vigilance of friends and foes in western lowland gorillas (Gorilla gorilla gorilla). Animal Behavior and Cognition, 7(4), 537-555. https://doi.org/10.26451/abc.07.04.06.2020
4. Kret, M.E., & Roth, T.S. (2020). Anecdotes in animal behaviour. Behaviour, 157(5), 385-386.
3. Roth, T.S., Rianti, P., Fredriksson, G., Wich, S.A., & Nowak, M.G. (2020). Grouping behavior of Sumatran orangutans (Pongo abelii) and Tapanuli orangutans (Pongo tapanuliensis) living in forest with low fruit abundance. American Journal of Primatology, 82(5), e23123. https://doi.org/10.1002/ajp.23123
2. Roth, T.S., Bionda, T.R., & Sterck, E.H.M. (2019). Duckling-catching by a zoo-housed western lowland gorilla (Gorilla gorilla gorilla). Journal of Zoo and Aquarium Research, 7(3), 144-148. https://doi.org/10.19227/jzar.v7i3.398
1. Roth, T.S., Bionda, T.R., & Sterck, E.H.M. (2017). Recapturing the canopy: stimulating Bornean orangutan (Pongo pygmaeus) locomotion behaviour in a zoo environment. Journal of Zoo and Aquarium Research, 5(1), 16–24. https://doi.org/10.19227/jzar.v5i1.186
Other:
Prüst, M., & Roth, T.S. (Eds.). 2014. 360° mens. Utrecht, NL: Uitgeverij de Graaff.

Publications under review
Bryon, E., Roth, T. S., Torfs, J. R. R., Eens, M., van Leeuwen, E. J. C, & Staes, N. Bonobos and chimpanzees are equally aggressive in within-group contexts yet differ in sex-specific ways. Under review in Science Advances.
Geijtenbeek, J. W., van den Berg, L. M., & Roth, T. S. Enclosures with flexible substrates stimulate species-typical locomotion, but not locomotion quantity, in captive Bornean orangutans (Pongo pygmaeus). Under review in Animal Welfare.
Samara, I., Fiacchino, D., Roth, T. S., de Vries, E., Kret, M. E., & Nikolic, M. The subtle art of seduction: Mimicry of coy smiles enhances interpersonal attraction. Under review in Behaviour.
Zijlstra, T. W., van Berlo, E., Roth, T. S., & Kret, M. E. Bonobos (Pan paniscus) increased attention to emotions may extend to human expressions. Under review in Scientific Reports