Scientific publications (see bottom of the page for manuscripts under review):
31. Roth, T. S., Massen, J. J. M., & Samara, I. (2026). Inconsistent Use of Evolutionary Frameworks in Accounts of Romantic Relationships (invited commentary). Behavioral and Brain Sciences, 49, e122. https://doi.org/10.1017/S0140525X25102331
30. Roth, T. S., Hirskyj-Douglas, I., & Perea-García, J. O. (2026). Designing Research Interfaces for Non-Human Minds: A Biocentric Design Approach for Animal Cognition Research. Animal Cognition. https://doi.org/10.1007/s10071-026-02069-x
29. Bhattacharjee, D., Zijlstra, T. W., Roth, T. S., Belli, E., Calis, S., Escriche Chova, P., Cousin, E., de Jong, J. A., De Laat, E. J. A. M., Guðjónsdóttir, A. R., Janmaat, K. R. L., Jeunick, E. J., Kluiver, C., Kuijer, P., Middelburg, E., Pflüger, L. S., Schroderus, V., van Dijk, E. S. J., Verspeek, J., Waasdorp, S., Zeeman, A., Sterck, E. H. M., van Leeuwen, E. J. C. & Massen, J.J.M. (2026). Despotism promotes cooperation through enhanced interdependencies in non-human primate societies. Nature Communications, 17, 3513. https://doi.org/10.1038/s41467-026-71168-7
28. Geijtenbeek, J. W., van den Berg, L. M., & Roth, T. S. (2026). Enclosures with flexible structures stimulate species-typical locomotion, but not locomotion quantity, in zoo-housed Bornean orangutans (Pongo pygmaeus). Animal Welfare, 35, e30. https://doi.org/10.1017/awf.2026.10086
27. Bryon, E., Roth, T. S., Torfs, J. R. R., Eens, M., van Leeuwen, E. J. C, & Staes, N. (2026). Bonobos and chimpanzees are equally aggressive in within-group contexts yet differ in sex-specific ways. Science Advances, 12(11), adz2433. https://doi.org/10.1126/sciadv.adz2433
26. Roth, T. S. (2026). Incorporating free partner choice into conservation breeding: the case for long-distance preference testing. Behaviour, https://doi.org/10.1163/1568539X-bja10346
25. van Leeuwen, E. J. C., & Roth, T. S. (2025). On the forces that bind us (invited commentary). Behavioral and Brain Sciences, 48, e186. https://doi.org/10.1017/S0140525X25101477
24. de Laat, E., Waasdorp, S., Roth, T. S., Massen, J. M. M., & Sterck, E. H. M. Despotic long-tailed macaques benefit others in a group service paradigm. Behaviour, 162(15), 1079–1107. https://doi.org/10.1163/1568539X-bja10338
23. Roth, T. S., van Berlo, E., Perea-García, J. O., & Kret, M. E. (2025). Bornean orangutans (Pongo pygmaeus) show an attentional bias toward a male secondary sexual trait. Annals of the New York Academy of Sciences, 1552, 225-238. http://doi.org/10.1111/nyas.70032
22. Sterck, E. H. M., Kamp, S. M., Rouart, I., van den Berg, L. M., Zijlmans, D. G. M., Roth, T. S., & Bushman, B. J. (2025). Reactions to Social Videos in Long-tailed Macaques. Animal Cognition, 28, 50. https://doi.org/10.1007/s10071-025-01970-1
21. van Oosten, J., Vanhooland, L. C., Roth, T. S., Gallup, A. C., Massen, J. M. M. (2025). No experimental evidence for yawn contagion in galahs (Eolophus roseicapilla). Animal Behavior & Cognition, 12(2), 148-171. https://doi.org/10.26451/abc.12.02.01.2025
20. Benítez, M. E., & Roth, T. S. (2024). Frans de Waal’s lasting legacy: Inspiring a new generation of animal behaviour scholars. Behaviour, 161(7), 511-522. https://doi.org/10.1163/1568539X-bja10277
19. Roth, T. S., von Wulffen, C., Fox, M., Askew, J., Spierings, M., & Perea-Garcia, J. O. (2024). Idiosyncratic long call-like vocalizations in two zoo-housed Bornean orang-utan females. Behaviour, 161(7), 567-585. https://doi.org/10.1163/1568539X-bja10273
18. Roth, T.S., Samara, I., Perea-Garcia, J.O., & Kret, M.E. (2024). No immediate attentional bias towards or choice bias for male secondary sexual characteristics in Bornean orang-utans (Pongo pygmaeus). Scientific Reports, 14, 12095. https://doi.org/10.1038/s41598-024-62187-9
17. Roth, T.S. (2024). Tinder for orang-utans: comparing sexually selective cognition among Bornean orang-utans (Pongo pygmaeus) and humans (Homo sapiens). PhD Thesis, Leiden University. https://hdl.handle.net/1887/3721951
16. Folz, J., Roth, T.S., Nikolic, M., & Kret, M.E. (2024). Who gets caught by the emotion? Attentional biases toward emotional facial expressions and their link to social anxiety and autistic traits. Current Psychology, 43, 3486–3502. https://doi.org/10.1007/s12144-023-04484-6
15. Roth, T.S., Samara, I., Perea-Garcia, J.O., & Kret, M.E. (2023). Individual attractiveness preferences differentially modulate immediate and voluntary attention. Scientific Reports, 13, 2147. https://doi.org/10.1038/s41598-023-29240-5
14. Laméris, D.W., van Berlo, E., Roth, T.S., & Kret, M.E. (2022). No evidence for biased attention towards emotional scenes in Bornean orangutans (Pongo pygmaeus). Affective Science, 3, 772-782. https://doi.org/10.1007/s42761-022-00158-x
13. Samara, I., Roth, T.S., Nikolic, M., Prochazkova, E., & Kret, M.E. (2022). Can third-party observers detect attraction in others based on subtle nonverbal cues? Current Psychology. https://doi.org/10.1007/s12144-022-02927-0
12. Sterck, E.H.M., & Roth, T.S. (2022). Socioecology of Asian colobines. In: The Colobines: Natural History, Behaviour, and Ecological Diversity (Matsuda I, Grueter C & Teichroeb JA, eds.). Cambridge University Press, UK (pp 250-270). https://doi.org/10.1017/9781108347150.017
11. Roth, T.S., Du, X., Samara, I., & Kret, M.E. (2022). Attractiveness modulates attention, but does not enhance gaze cueing. Evolutionary Behavioral Sciences, 16(4), 343-361. https://doi.apa.org/doi/10.1037/ebs0000265
10. Samara, I., Roth, T.S., & Kret, M.E. (2021). The Role of Emotion Projection, Sexual Desire, and Self-Rated Attractiveness in the Sexual Overperception Bias. Archives of Sexual Behavior, 50, 2507–2516. https://doi.org/10.1007/s10508-021-02017-5
9. Nikolic, M., & Roth, T.S. (2021). A comparative glance on self-conscious emotions: A commentary on Kret et al. (2020). Neuroscience & Biobehavioral Reviews, 129, 154-156. https://doi.org/10.1016/j.neubiorev.2021.07.031
8. Roth, T.S., Samara, I., & Kret, M.E. (2021). Multimodal mate choice: Exploring the effects of sight, sound, and scent on partner choice in a speed-date paradigm. Evolution and Human Behavior, 42(5), 461-468. https://doi.org/10.1016/j.evolhumbehav.2021.04.004
7. Roth, T.S., Samara, I., Tan, J., Prochazkova, E., & Kret, M.E. 2021. A comparative framework of inter-individual coordination and pair-bonding. Current Opinion in Behavioral Sciences 39. https://doi.org/10.1016/j.cobeha.2021.03.005
6. Roth, T.S., Samara, I., & Kret, M.E. (2021). Ultimate and proximate factors underlying sexual overperception bias: A reply to Lee et al. (2020). Evolution and Human Behavior, 40(1), 73-75. https://doi.org/10.1016/j.evolhumbehav.2020.06.002
5. Roth, T.S., & Sterck, E.H.M. (2020). Social vigilance of friends and foes in western lowland gorillas (Gorilla gorilla gorilla). Animal Behavior and Cognition, 7(4), 537-555. https://doi.org/10.26451/abc.07.04.06.2020
4. Kret, M.E., & Roth, T.S. (2020). Anecdotes in animal behaviour. Behaviour, 157(5), 385-386.
3. Roth, T.S., Rianti, P., Fredriksson, G., Wich, S.A., & Nowak, M.G. (2020). Grouping behavior of Sumatran orangutans (Pongo abelii) and Tapanuli orangutans (Pongo tapanuliensis) living in forest with low fruit abundance. American Journal of Primatology, 82(5), e23123. https://doi.org/10.1002/ajp.23123
2. Roth, T.S., Bionda, T.R., & Sterck, E.H.M. (2019). Duckling-catching by a zoo-housed western lowland gorilla (Gorilla gorilla gorilla). Journal of Zoo and Aquarium Research, 7(3), 144-148. https://doi.org/10.19227/jzar.v7i3.398
1. Roth, T.S., Bionda, T.R., & Sterck, E.H.M. (2017). Recapturing the canopy: stimulating Bornean orangutan (Pongo pygmaeus) locomotion behaviour in a zoo environment. Journal of Zoo and Aquarium Research, 5(1), 16–24. https://doi.org/10.19227/jzar.v5i1.186
Other:
Prüst, M., & Roth, T.S. (Eds.). 2014. 360° mens. Utrecht, NL: Uitgeverij de Graaff.
Publications under review
Samara, I., Fiacchino, D., Roth, T. S., de Vries, E., Kret, M. E., & Nikolic, M. The subtle art of seduction: Mimicry of coy smiles enhances interpersonal attraction. Under review in Behaviour.
Zijlstra, T. W., van Berlo, E., Roth, T. S., & Kret, M. E. Bonobos (Pan paniscus) increased attention to emotions may extend to human expressions. Under review in Scientific Reports